Nathan C. Fillion was born on March 27, 1971 in Edmonton, Alberta, Canada. He is a Canadian actor.

He proved very popular as the show’s character Joey Buchanan and was nominated for a Daytime Emmy Award, as well as a Soap Opera Digest Award, for “Outstanding Younger Actor” in 1996.

After three years in the role, on the advice of a mentor, Fillion moved to Los Angeles to persue further career opportunities. The move proved fruitful and gained him roles in the films Saving Private Ryan, Blast From the Past, and Dracula 2000.

A recurring guest role on Two Guys and a Girl as the lovable Johnny Donnelly was so popular the character become a full time role. When Two Guys and a Girl ended, Fox gave Fillion a talent holding deal which lead to the leading role in Firefly in 2003.

Fillion proved that his likability is not a hindrance to playing a villain when he was cast in the final five episodes of Buffy the Vampire Slayer in 2003. Delivering deliciously evil one-liners with a smile and a gleam in his eye, while effortlessly snapping the necks of “dirty girls,” Fillion thoroughly stole his scenes as the sweet and mannerly, yet horrific female-hating preacher, Caleb.

He received critical praise for his roles in two independent Canadian films, Water’s Edge (2003) and Outing Riley (2004). Playing grieving novelist Robert Graves in Water’s Edge, Fillion demonstrated his flair for subtle realism. As family jokester Luke Riley in Outing Riley, his ability to combine comedy with sensitive dramatic material was beautifully revealed.

Serenity was followed up by the leading role in the quirky horror film, Slither (2006), which won him glowing reviews across the board for his understated performance as the hapless, frustrated hero, Police Chief Bill Pardy.

Other roles include the indie film Waitress (2007), White Noise 2: The Light (2007), horror flick Slither (2006) and TV series "Desperate Housewives" (2007-2008).

Filmography

• Chilled in Miami (2009)
• Trucker (2008)
• "Desperate Housewives" (8 episodes, 2007-2008)
• "Robot Chicken" (1 episode, 2007)
• "One Life to Live" (2 episodes, 1994-2007)
• "Drive" (6 episodes, 2007)
• Waitress (2007)
• White Noise 2: The Light (2007)
• "Lost" (1 episode, 2006)
• Slither (2006)
• "Justice League" (2 episodes, 2005-2006)
• Serenity (2005)
• "Miss Match" (6 episodes, 2003-2004)
• Outing Riley (2004)
• Hollywood Division (2004) (TV)
• Water's Edge (2003)
• "Firefly" (14 episodes, 2002-2003)
• "Buffy the Vampire Slayer" (5 episodes, 2003)
• Alligator Point (2003) (TV)
• "Pasadena" (3 episodes, 2002)
• "Two Guys, a Girl and a Pizza Place" (53 episodes, 1998-2001)
• "King of the Hill" (1 episode, 2001)
• Dracula 2000 (2000)
• "The Outer Limits" (1 episode, 1999)
• Blast from the Past (1999)
• "Maggie Winters" (1 episode, 1998)
• Saving Private Ryan (1998)
• "Total Security" (1 episode 1997)
• "Spin City" (1 episode, 1996)
• Strange and Rich (1994)
• Ordeal in the Arctic (1993) (TV)